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Biology — Heritability

Narrow- and broad-sense heritability.

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h^2 = \frac{V_A}{V_P} \;\text{(narrow sense)}

H^2 = \frac{V_G}{V_P} \;\text{(broad sense)}

V_P = V_G + V_E

R = h^2 S \;\text{(response to selection)}