Biology — Selection & Mean Fitness
Mean population fitness and the change in allele frequency.
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\bar{w} = p^2 w_{AA} + 2pq\,w_{Aa} + q^2 w_{aa}
p' = \frac{p^2 w_{AA} + pq\,w_{Aa}}{\bar{w}}
\Delta p = p' - p
w_i = 1 - s_i